Trilobite fossils
Trilobites
Trilobites ("three-lobes")
are extinct arthropods that form the class Trilobita.
They appeared in the Middle Cambrian epoch and flourished throughout
the lower Paleozoic era before beginning a drawn-out decline
to extinction when, during the Late Devonian extinction, all
trilobite orders, with the sole exception of Proetida, died out.
The last of the trilobites disappeared in the mass extinction
at the end of the Permian about 250 million years ago.
Trilobite fossils are very well-known, and possibly the second-most
famous fossil group, after the dinosaurs. When trilobites appear
in the fossil record of the Lower Cambrian they are already highly
diverse and geographically dispersed. Because of their diversity
and an easily fossilized exoskeleton, they left an extensive fossil
record with some 17,000 known species spanning Paleozoic time.
Trilobite fossils have been important in biostratigraphy, paleontology,
and plate tectonics research. For example, trilobite fossils have
been important in estimating the rate of speciation during the
period known as the Cambrian Explosion because they are the most
diverse group of metazoans known from the fossil record of the
early Cambrian (Lieberman, 1999), and are readily distinguishable
because of complex and well preserved morphologies. The trilobites
are often placed within the arthropod subphylum Schizoramia within
the superclass Arachnomorpha (equivalent to the Arachnata) (e.g.,
Cotton & Braddy 2004), although several alternative taxonomies
are found in the literature.
Different trilobites made their living in different ways. Some
led a benthic life as predators, scavengers or filter feeders.
Some swam (a pelagic lifestyle) and fed on plankton. Still others
(particularly the family Olenidae) are thought to have evolved
a symbiotic relationship with sulfur-eating bacteria from which
they derived food.
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