Psittacosaurus fossils
Psittacosaurus
Psittacosaurus is a genus of psittacosaurid
ceratopsian dinosaur from the Early Cretaceous Period of what is
now Asia. It is notable for being the most species-rich dinosaur
genus. At least ten extinct species are recognized from dinosaur
fossils found in different regions of modern-day China, Mongolia
and Russia.
All species of Psittacosaurus were gazelle-sized bipedal herbivores
characterized by a high, powerful beak on the upper jaw. At least one species
had long, quill-like structures on its tail and lower back, however recent research
suggests these may not be quills or bristles but rather collagen fibrils. Psittacosaurs
were extremely early ceratopsians and, while they developed many novel adaptations
of their own, they also shared many anatomical features with later ceratopsians,
such as Protoceratops and Triceratops dinosaurs.
Psittacosaurus is not as familiar to the general public as its distant
relative Triceratops but it is one of the most completely known dinosaur
genera. Dinosaur bones of over 400 individual dinosaur skeletons have been collected
so far, including many complete fossil skeletons. Most different age classes
are represented, from hatchling through to adult, which has allowed several detailed
studies of Psittacosaurus growth rates and reproductive biology. The
abundance of this dinosaur in the fossil record has led to its use as an index
fossil for Early Cretaceous sediments of central Asia.
Different species of Psittacosaurus varied in size and specific features
of the dinosaur skull and dinosaur skeleton, but shared the same overall body
shape. The best-known species, P. mongoliensis, reached 2 metres
(6.5 ft) in length. The maximum adult body weight was most likely over 20 kilograms
(44 lb) in P. mongoliensis. Several species approached P. mongoliensis in
size (P. major, P. neimongoliensis, P. xinjiangensis),
while others were somewhat smaller (P. sinensis, P. meileyingensis). P.
ordosensis was the smallest known species, 30% smaller than P. mongoliensis.
The largest were P. lujiatunensis and P. sibiricus, although
neither was significantly larger than P. mongoliensis.
The skull of Psittacosaurus was highly modified compared to other ornithischian
dinosaurs of its time. The dinosaurs skull was extremely tall and short, with
an almost round profile in some species. The portion in front of the orbit (eye
socket) was only 40% of total skull length, shorter than any other known ornithischian.
The lower jaws of psittacosaurs are characterized by a bulbous vertical ridge
down the center of each tooth. Both upper and lower jaws sported a pronounced
beak, formed from the rostral and predentary bones, respectively.
The bony core of the beak may have been sheathed in keratin to provide a sharp
cutting surface for cropping plant material. As the generic name suggests, the
short skull and beak superficially resembled those of modern parrots. Psittacosaurus skulls
shared several adaptations with more derived ceratopsians, such as the unique
rostral bone at the tip of the upper jaw, and the flared jugal (cheek) bones.
However, there was still no sign of the bony neck frill or prominent facial horns
which would develop in later ceratopsians.
Psittacosaurus postcranial dinosaur skeletons were more typical of a
'generic' bipedal ornithischian. In P. mongoliensis, similarly to other
species, the forelimbs were only 58% as long as the hindlimbs, indicating that
these animals were almost totally bipedal in life. There were only four digits
on the manus ('hand'), as opposed to the five found in most other ornithischians
(including all other ceratopsians). Overall, the four-toed hindfoot was very
similar to many other small ornithischians.
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